What resolves this contention: Anatomical extent of the layer-1 thalamic input — restricted matrix population (Jones-style) vs broad, dense canopy contributed by most thalamic nuclei (Rubio-Garrido). / Layer-1-projecting thalamic axons originate from most rodent thalamic nuclei and form a dense, highly convergent thalamic canopy on apical tufts (~4500 cells/mm²). / Layer-1 thalamocortical projections come predominantly from a calbindin matrix in selected nuclei and provide diffuse, modulatory rather than dense convergent input.
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