We will apply Dirichlet regression with Braak stage as the primary predictor to SEA-AD snRNA-seq subtype proportions (84 donors, DLPFC) and replicate in ROSMAP (Mathys 2023, ~427 donors). For each major microglia subtype (homeostatic MG-1, transitional MG-2, DAM/MGnD MG-3, MG-4) and each oligodendrocyte/OPC subtype, we will estimate Braak-stage inflection points using piecewise-linear or segmented regression with bootstrap 95% CIs. Cross-lineage comparison will test whether the microglia inflection precedes, follows, or overlaps the oligo/OPC inflection (non-overlapping CIs as the primary criterion). Sex, APOE genotype, and post-mortem interval will enter as covariates. Replication in ROSMAP and directional consistency with Green et al. 2024 cellular communities will be required before causal language is used.

Details

disease
alzheimer's disease
target_ref
dataset:SEA-AD-snRNA-DLPFC
identification_strategy
observational
Raw fields (4)
assay_spec
Primary cohort: SEA-AD snRNA-seq DLPFC, 84 donors, accessed via CELLxGENE Census Python API (cellxgene_census.get_obs, tissue_general='prefrontal cortex', dataset_title contains 'SEA-AD'). Microglia subtypes resolved from obs field 'subclass_label' (homeostatic: P2RY12+/TMEM119+; DAM/MGnD: SPP1+/GPNMB+/TREM2+; IRM: IFIT1+/IFIT3+; LDA: LPL+/PLIN2+). OPC and oligodendrocyte subtypes from same field (OPC: PDGFRA+; mature oligo: MBP+/MOG+). Predictor: braak_int (0–6, mapped from Braak_stage obs field). Covariates: sex, PMI, RIN, age_at_death. Replication cohort: Mathys 2023 Cell (DOI: 10.1016/j.cell.2023.08.039), GEO GSE174367, per-donor proportion table from supplement. Independent validation: Green 2024 Nature (DOI: 10.1038/s41586-024-07871-6) ROSMAP cellular-community fractions. Statistical model: scCODA (pertpy>=0.7 Sccoda), num_samples=10000, num_warmup=2000, credible_effects_threshold=0.95. Planned output: inflection-point Braak stage per subtype (posterior effect sign reversal or credible threshold crossing), reported with 95% HDI.
hypothesis
DAM/MGnD microglia (SPP1+, GPNMB+, TREM2+) proportions rise at an earlier Braak stage than oligodendrocyte/OPC proportions fall, indicating that neuroinflammatory expansion precedes—and may drive—oligodendroglial depletion in AD cortex rather than the reverse.
kill_criteria
Abort or revise if: (1) CELLxGENE Census obs lacks donor_id or Braak_stage — BLOCKED, file Allen portal data-request; (2) R-hat > 1.05 on any posterior parameter in microglia or oligo model — flag, withhold interpretation; (3) Mathys 2023 GEO supplement lacks per-donor count matrix with Braak metadata — replication arm BLOCKED, fall back to Mathys 2019 or report single-cohort only; (4) fewer than 5 donors per Braak stage bin in SEA-AD — compositional model underpowered, report with caveat; (5) Green 2024 community fractions do not map to microglia or oligo subtypes — validation arm BLOCKED; (6) logistic curve_fit fails to converge for >50% of subtypes — inflection detection BLOCKED, use empirical mode instead; (7) DAM/MGnD posterior HDI overlaps OPC/oligo nadir HDI across all Braak bins — hypothesis NOT supported, revise to null-result artifact.
primary_endpoint
Tick 12: Targeted EuropePMC sweeps with shorter query strings — (1) microglia DAM Braak snRNA human, (2) OPC/oligodendrocyte depletion AD cortex snRNA — plus CrossRef provenance stamps for Mathys 2019 (DOI 10.1038/s41586-019-1195-2) and Mathys 2023 (DOI 10.1016/j.cell.2023.08.039) as ROSMAP anchor references. Success criterion: ≥1 hit per sweep with human snRNA-seq microglia subtype or OPC staging data; if both criteria met, advance to notebook scaffold tick.

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