What resolves this contention: Whether top-down vs neuromodulatory inputs are t…

What resolves this contention: Whether top-down vs neuromodulatory inputs are the primary driver of cortical VIP recruitment / VIP recruitment during arousal/locomotion in V1 is driven primarily by cholinergic basal-forebrain inputs via nicotinic receptors (state/neuromodulatory). / VIP recruitment in S1 and during attention is driven primarily by long-range glutamatergic top-down inputs from frontal/secondary motor areas (cingulate, vM1).