Computational Models of VIP Circuit Function

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Computational Models of VIP Circuit Function

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Source: https://github.com/AllenNeuralDynamics/ComputationalReviewVIP/blob/95e761177f7d2ec565983d3307c14ec238f9677c/content/12_computational_models.md

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  • 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...

  • 2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...

  • 3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...

  • 4Citationpaper:paper-5cfe39bf225dComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...

  • 5Citationpaper:paper-acf576c0a2edComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...

  • 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference0 Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference1; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...

  • 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2 Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...

  • 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference4 Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference5; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...

  • 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference6 Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference7; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...

  • 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference8 Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference9; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...

  • 2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference0 Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output 2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference1; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...

  • 2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2 Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output 2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...

  • 2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference4 Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output 2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference5; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...

  • 2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference6 Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output 2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference7; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...

  • 2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference8 Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output 2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference9; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...

  • 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference0 Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference1; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...

  • 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2 Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...

  • 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference4 Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference5; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...

  • 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference6 Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference7; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...

  • 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference8 Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference9; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...

  • 3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference0 Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output 3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference1; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...

  • 3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2 Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output 3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...

  • 3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference4 Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output 3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference5; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...

  • 3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference6 Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output 3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference7; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...

  • 3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference8 Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output 3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference9; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...

  • 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference0 Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference1; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...

  • 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2 Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...

  • 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference4 Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference5; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...

  • 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference6 Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference7; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...

  • 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference8 Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference9; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...

  • 4Citationpaper:paper-5cfe39bf225dComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference0 Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output 4Citationpaper:paper-5cfe39bf225dComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference1; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...

  • 4Citationpaper:paper-5cfe39bf225dComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2 Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output 4Citationpaper:paper-5cfe39bf225dComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...

  • 4Citationpaper:paper-5cfe39bf225dComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference4 Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output 4Citationpaper:paper-5cfe39bf225dComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference5; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...

  • 4Citationpaper:paper-5cfe39bf225dComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference6 Standard three-interneuron (3-IN) circuits — Pyr+PV+SST+VIP — emerged from the connectivity surveys of 4Citationpaper:paper-5cfe39bf225dComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference7 and the disinhibitory experiments of 4Citationpaper:paper-5cfe39bf225dComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference8, and have been formalised as rate models 4Citationpaper:paper-5cfe39bf225dComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference9, spiking implementations 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference0, and biophysically detailed mean-field circuits 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference1. Across this family, VIP→SST di...

  • 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2 Standard three-interneuron (3-IN) circuits — Pyr+PV+SST+VIP — emerged from the connectivity surveys of 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3 and the disinhibitory experiments of 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference4, and have been formalised as rate models 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference5, spiking implementations 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference6, and biophysically detailed mean-field circuits 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference7. Across this family, VIP→SST di...

  • 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference8 Standard three-interneuron (3-IN) circuits — Pyr+PV+SST+VIP — emerged from the connectivity surveys of 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference9 and the disinhibitory experiments of 5Citationpaper:paper-acf576c0a2edComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference0, and have been formalised as rate models 5Citationpaper:paper-acf576c0a2edComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference1, spiking implementations 5Citationpaper:paper-acf576c0a2edComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2, and biophysically detailed mean-field circuits 5Citationpaper:paper-acf576c0a2edComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3. Across this family, VIP→SST di...

  • 5Citationpaper:paper-acf576c0a2edComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference4 Standard three-interneuron (3-IN) circuits — Pyr+PV+SST+VIP — emerged from the connectivity surveys of 5Citationpaper:paper-acf576c0a2edComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference5 and the disinhibitory experiments of 5Citationpaper:paper-acf576c0a2edComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference6, and have been formalised as rate models 5Citationpaper:paper-acf576c0a2edComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference7, spiking implementations 5Citationpaper:paper-acf576c0a2edComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference8, and biophysically detailed mean-field circuits 5Citationpaper:paper-acf576c0a2edComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference9. Across this family, VIP→SST di...

  • 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference0 Standard three-interneuron (3-IN) circuits — Pyr+PV+SST+VIP — emerged from the connectivity surveys of 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference1 and the disinhibitory experiments of 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2, and have been formalised as rate models 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3, spiking implementations 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference4, and biophysically detailed mean-field circuits 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference5. Across this family, VIP→SST di...

  • 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference6 Standard three-interneuron (3-IN) circuits — Pyr+PV+SST+VIP — emerged from the connectivity surveys of 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference7 and the disinhibitory experiments of 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference8, and have been formalised as rate models 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference9, spiking implementations 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference00, and biophysically detailed mean-field circuits 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference01. Across this family, VIP→SST di...

  • 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference02 Standard three-interneuron (3-IN) circuits — Pyr+PV+SST+VIP — emerged from the connectivity surveys of 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference03 and the disinhibitory experiments of 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference04, and have been formalised as rate models 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference05, spiking implementations 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference06, and biophysically detailed mean-field circuits 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference07. Across this family, VIP→SST di...

  • 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference08 Standard three-interneuron (3-IN) circuits — Pyr+PV+SST+VIP — emerged from the connectivity surveys of 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference09 and the disinhibitory experiments of 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference10, and have been formalised as rate models 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference11, spiking implementations 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference12, and biophysically detailed mean-field circuits 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference13. Across this family, VIP→SST di...

  • 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference14 Standard three-interneuron (3-IN) circuits — Pyr+PV+SST+VIP — emerged from the connectivity surveys of 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference15 and the disinhibitory experiments of 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference16, and have been formalised as rate models 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference17, spiking implementations 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference18, and biophysically detailed mean-field circuits 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference19. Across this family, VIP→SST di...

  • 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference20 Standard three-interneuron (3-IN) circuits — Pyr+PV+SST+VIP — emerged from the connectivity surveys of 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference21 and the disinhibitory experiments of 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference22, and have been formalised as rate models 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference23, spiking implementations 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference24, and biophysically detailed mean-field circuits 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference25. Across this family, VIP→SST di...

  • 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference26 Standard three-interneuron (3-IN) circuits — Pyr+PV+SST+VIP — emerged from the connectivity surveys of 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference27 and the disinhibitory experiments of 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference28, and have been formalised as rate models 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference29, spiking implementations 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference30, and biophysically detailed mean-field circuits 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference31. Across this family, VIP→SST di...

  • 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference32 Standard three-interneuron (3-IN) circuits — Pyr+PV+SST+VIP — emerged from the connectivity surveys of 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference33 and the disinhibitory experiments of 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference34, and have been formalised as rate models 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference35, spiking implementations 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference36, and biophysically detailed mean-field circuits 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference37. Across this family, VIP→SST di...

  • 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference38 Standard three-interneuron (3-IN) circuits — Pyr+PV+SST+VIP — emerged from the connectivity surveys of 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference39 and the disinhibitory experiments of 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference40, and have been formalised as rate models 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference41, spiking implementations 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference42, and biophysically detailed mean-field circuits 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference43. Across this family, VIP→SST di...

  • 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference44 Standard three-interneuron (3-IN) circuits — Pyr+PV+SST+VIP — emerged from the connectivity surveys of 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference45 and the disinhibitory experiments of 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference46, and have been formalised as rate models 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference47, spiking implementations 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference48, and biophysically detailed mean-field circuits 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference49. Across this family, VIP→SST di...

  • 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference50 Standard three-interneuron (3-IN) circuits — Pyr+PV+SST+VIP — emerged from the connectivity surveys of 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference51 and the disinhibitory experiments of 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference52, and have been formalised as rate models 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference53, spiking implementations 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference54, and biophysically detailed mean-field circuits 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference55. Across this family, VIP→SST di...

  • 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference56 Standard three-interneuron (3-IN) circuits — Pyr+PV+SST+VIP — emerged from the connectivity surveys of 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference57 and the disinhibitory experiments of 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference58, and have been formalised as rate models 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference59, spiking implementations 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference60, and biophysically detailed mean-field circuits 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference61. Across this family, VIP→SST di...

  • 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference62 Standard three-interneuron (3-IN) circuits — Pyr+PV+SST+VIP — emerged from the connectivity surveys of 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference63 and the disinhibitory experiments of 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference64, and have been formalised as rate models 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference65, spiking implementations 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference66, and biophysically detailed mean-field circuits 1Citationpaper:paper-065a0eae69a5Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference2Citationpaper:paper-1659c2dc5d60Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference3Citationpaper:paper-f02f24cb44ffComputational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...content/12_computational_models.md:line 5Open reference67. Across this family, VIP→SST di...

  • ... 345 additional anchors in refs_json

References

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  2. [Pfeffer2013] paper:paper-1659c2dc5d60 “Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...”
  3. [Lee2013] paper:paper-f02f24cb44ff “Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...”
  4. [Fu2014] paper:paper-5cfe39bf225d “Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...”
  5. [Pakan2016] paper:paper-acf576c0a2ed “Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...”
  6. [Krabbe2019] paper:paper-6864c305085a “Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...”
  7. [Dipoppa2018] paper:paper-f457e091013f “Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...”
  8. [Garciadelmolino2017] paper:paper-e67fae66877b “Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...”
  9. [Litwinkumar2016] paper:paper-8f25714ab7fe “Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...”
  10. [Hertag2019] paper:paper-485852ecd519 “Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...”
  11. [Bos2025] paper:paper-00de1a2c593e “Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...”
  12. [Veit2023] paper:paper-d895aafe5d83 “Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...”
  13. [Iqbal2025] paper:paper-733c0535ffcd “Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...”
  14. [Hartung2024] paper:paper-41c66384d08f “Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...”
  15. [Anastasiades2021a] paper:paper-7227d2a3ed3e “Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...”
  16. [Lee2025] paper:paper-193dfb3c1e40 “Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...”
  17. [Hertag2020] paper:paper-3a51e8e844f7 “Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...”
  18. [Hertag2021] paper:paper-ab222bc802a5 “Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...”
  19. [Rossbroich2025] paper:paper-e82e733c3a59 “Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...”
  20. [Nemati2025a] paper:paper-1eeff23c6853 “Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...”
  21. [Nemati2025b] paper:paper-d67d30eba36a “Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...”
  22. [Shipp2016] paper:paper-1a025491591d “Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...”
  23. [Chevy2024] paper:paper-128b1d42ab7b “Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...”
  24. [Shen2023] paper:paper-d557fa86de56 “Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...”
  25. [Yang2016] paper:paper-1bdad28ffcad “Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...”
  26. [Tahvili2025a] paper:paper-fe7e8b76572a “Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...”
  27. [Tahvili2025b] paper:paper-2e1f55b5f880 “Computational models of cortex have, over the past decade, moved VIP interneurons from a peripheral curiosity to an indispensable structural element. Three pressures forced this shift: the demonstration that VIP cells preferentially target SST cells and disinhibit pyramidal output [Pi2013,Pfeffer2013,Lee2013]; the discovery that VIP firing tracks state, locomotion, reward, and reinforcement signals [Fu2014,Pakan2016...”
  28. [Reimann2026] paper:paper-aedf4d3b8990 “Standard three-interneuron (3-IN) circuits — Pyr+PV+SST+VIP — emerged from the connectivity surveys of [Pfeffer2013] and the disinhibitory experiments of [Pi2013,Lee2013,Fu2014], and have been formalised as rate models [Litwinkumar2016,Garciadelmolino2017,Hertag2019], spiking implementations [Veit2023], and biophysically detailed mean-field circuits [Reimann2026,Jadi2012,Isbister2026]. Across this family, VIP→SST di...”
  29. [Jadi2012] paper:paper-1b029487a634 “Standard three-interneuron (3-IN) circuits — Pyr+PV+SST+VIP — emerged from the connectivity surveys of [Pfeffer2013] and the disinhibitory experiments of [Pi2013,Lee2013,Fu2014], and have been formalised as rate models [Litwinkumar2016,Garciadelmolino2017,Hertag2019], spiking implementations [Veit2023], and biophysically detailed mean-field circuits [Reimann2026,Jadi2012,Isbister2026]. Across this family, VIP→SST di...”
  30. [Isbister2026] paper:paper-3f260fc5a348 “Standard three-interneuron (3-IN) circuits — Pyr+PV+SST+VIP — emerged from the connectivity surveys of [Pfeffer2013] and the disinhibitory experiments of [Pi2013,Lee2013,Fu2014], and have been formalised as rate models [Litwinkumar2016,Garciadelmolino2017,Hertag2019], spiking implementations [Veit2023], and biophysically detailed mean-field circuits [Reimann2026,Jadi2012,Isbister2026]. Across this family, VIP→SST di...”
  31. [Karnani2016a] paper:paper-d405e5906927 “Standard three-interneuron (3-IN) circuits — Pyr+PV+SST+VIP — emerged from the connectivity surveys of [Pfeffer2013] and the disinhibitory experiments of [Pi2013,Lee2013,Fu2014], and have been formalised as rate models [Litwinkumar2016,Garciadelmolino2017,Hertag2019], spiking implementations [Veit2023], and biophysically detailed mean-field circuits [Reimann2026,Jadi2012,Isbister2026]. Across this family, VIP→SST di...”
  32. [Abs2018] paper:paper-cddfd665bb4c “Standard three-interneuron (3-IN) circuits — Pyr+PV+SST+VIP — emerged from the connectivity surveys of [Pfeffer2013] and the disinhibitory experiments of [Pi2013,Lee2013,Fu2014], and have been formalised as rate models [Litwinkumar2016,Garciadelmolino2017,Hertag2019], spiking implementations [Veit2023], and biophysically detailed mean-field circuits [Reimann2026,Jadi2012,Isbister2026]. Across this family, VIP→SST di...”
  33. [Hahn2020] paper:paper-a201ea93b617 “Standard three-interneuron (3-IN) circuits — Pyr+PV+SST+VIP — emerged from the connectivity surveys of [Pfeffer2013] and the disinhibitory experiments of [Pi2013,Lee2013,Fu2014], and have been formalised as rate models [Litwinkumar2016,Garciadelmolino2017,Hertag2019], spiking implementations [Veit2023], and biophysically detailed mean-field circuits [Reimann2026,Jadi2012,Isbister2026]. Across this family, VIP→SST di...”
  34. [Kim2025b] paper:paper-732f0f33eade “Standard three-interneuron (3-IN) circuits — Pyr+PV+SST+VIP — emerged from the connectivity surveys of [Pfeffer2013] and the disinhibitory experiments of [Pi2013,Lee2013,Fu2014], and have been formalised as rate models [Litwinkumar2016,Garciadelmolino2017,Hertag2019], spiking implementations [Veit2023], and biophysically detailed mean-field circuits [Reimann2026,Jadi2012,Isbister2026]. Across this family, VIP→SST di...”
  35. [Park2020] paper:paper-72a15d353d01 “Standard three-interneuron (3-IN) circuits — Pyr+PV+SST+VIP — emerged from the connectivity surveys of [Pfeffer2013] and the disinhibitory experiments of [Pi2013,Lee2013,Fu2014], and have been formalised as rate models [Litwinkumar2016,Garciadelmolino2017,Hertag2019], spiking implementations [Veit2023], and biophysically detailed mean-field circuits [Reimann2026,Jadi2012,Isbister2026]. Across this family, VIP→SST di...”
  36. [Natan2017] paper:paper-542ae9c3435e “Standard three-interneuron (3-IN) circuits — Pyr+PV+SST+VIP — emerged from the connectivity surveys of [Pfeffer2013] and the disinhibitory experiments of [Pi2013,Lee2013,Fu2014], and have been formalised as rate models [Litwinkumar2016,Garciadelmolino2017,Hertag2019], spiking implementations [Veit2023], and biophysically detailed mean-field circuits [Reimann2026,Jadi2012,Isbister2026]. Across this family, VIP→SST di...”

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